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Abstract The present investigation was undertaken at the Faculty of Agriculture of Moshtohor (zagazig University Benha Branch)’ The objective of this investigation waS concerned with genetic and cytological studies on Q~~21!:!~.!:!! spp found in Egypt COffi’p’8reao n wi th some e x o t i c S ~f l”OITlt.: Germany.’ Stu d~te 5 Inc 1u dec’ f0 u r sp eC 1t:: ~ 0 f the ge nus Oc not her a ,-; i.e.jQ. !l.i.~~~~~~ v a r fiedled, Q. ~l~~~~ L. var. grandiflora, Q. ~2~~~i[er!! renner and Q. £~~~~~~ jacy. Karyotype studies were carried out on these species at the mitotic metaphase stage. All species studied had the same chromosome basic numbcr (x = 7) in the smoatic cell. These chromosomes were metacentric, sub-mctacentric, acrocentric and subacrocentric. The following characters were studi ed 1- Earliness. 2- Plant heLg~t 3- Number of fruits per plan t 4- ~~I!’bcr of seeds per f r u i t 5- Weight of seeds per plant 6- 1000- seeds we i.ght R-e-s-u-l-t-s---s-h-o-we-d---t-h-a-t-- (A): Additive genetic variances were more important than dominance in the inheritance of Earlines. Dominance genetic variances were more important than additive in the inheretance of p la n t height,and numbe r of fruit’! plant fn the inheritance of weight seed per plant and weight of 100 seeds The add i t ive and dom ina n c e t y pes ) 0 f g e n e act ion. were imp 0r tan tin the inher ita nceo f;:the sc .tr a its . (B): The average degree of dominance showed over complete dominance whil~ partial dominanc number of f~uit/ pl&nt. showed showed dominance for plant height Earl i ne ss (C): The negative and positive alleles were equally distributed among the pare~tal populations for most of the studied characters. (D): The dominant genes were in excess in the parents in case of plant height and number of fruit/plant and weight of seed/plant and weight of IOOO-seeds. The values of narrow sense heritability in all characters indicated that the traits were highly influenced by the genetic factors. There were positive genotypic correlations between earliness and plant height and weight of seed! plant and we igh tor 1000 seed sand numbe r 0 f f ru it.S I plan 1. There were positive genotypic correlation between plant height and the number of seeds! fruit. Pi rst: Mi totic behaviour: studies included the : I Kar, otype and Ka~~)otype analys i s for four parent s 3 • The following description summariz~ the characteristics of the studied material 1- o. nissensis: 2n14. The average total length of the hoploid complement in the five cells studied was 29.10 + 0.41~. The mean length of the chromosemes ranged from 0.95 to 5.84 p. The idiogram showed that the 6th and 7th chromosomes were subacrocentric. 2- Q: biennis: 2n= 14. The average total length of the haploid complement was 25.45 + 0.12 p. The mean length of the chromosomes ranged from 1.25 to 7.06 p. The idiogram showed that the 3!~ chromosome was acrocentic with a small satellite attachl~d_ to its short arm. 3- o. coronifera: 2n= 14. The average total length of the haploid complement was 30.22 + 0.06 p. The mean length of the chromosomes ranged from 2.06 to The idio gram showed that the 6th chromosome was subacrocentric. 4- o. odorata: 2n= 14. The average length of the haploid complement was 19.00 + 0.08 p. The mean length of the chr~osomes ranged from 1.51 to 4.49 p. The idiogram showed that the 5!~, 6th and 7th chromosomes were subacrocentric. Second: Miotic behaviour’ These studies included the a b nor rnalit ies 0 f par e n tie san d the hy brid s P r0 ducE: c b ’f d ia Ll e crosses, it. was :not’ic’ed that ther,e are~some c@s~eS o f rin g s • and zig zag e w hie 11 : rnayin d icat e the v reS en ce ’ 0 f c e r t ~ n .Ch r ono sane tr arr s:10 ea tl ons , In these studies the pollen grain mother cells were examined in the stage of metaphase two (diakinases). ~he StudY showed the following: r: l h.1<;: a- ’fhc chromosomes appeared as circles consisting of four five or sex chromosomes. b- There were group conststing of two or three or four five chromosomes in a zigzag shape. c- There were chromosomes in star shape or a linear shape or in bivalents. e- There were obend rings of chromosomes. There were br idges chromosomes, and 1_~_gLn_g chromosomes ins orne cas e s 0 f par en t e san d h Y b rid s . W nic h mt:\ ’/ in d i.cat e dthe Pi’G’>~ ;le,~ )f some acrocentric from t r.a n s 10 c t ion s • 3) POll~n grain viability was high in some hybrids and low in others this strility also, i could be a”t r i bu t e dhet e r 0 z.y’-gou·s t 0 c eft a inc h r orno s om a 1 abe r rat ion sin c led i ng tranlocation |